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That’s not true. Only 26% (and shrinking) of Israeli Jews have European ancestry. A plurality, 44% (and the fastest growing) have native Israeli ancestry. The remainder come from North Africa and Asian ancestry but all groups besides Israeli background are shrinking.
The study you're referencing is looking at paternal lineage of Israelis born in Israel, not the ancestral lineage of all Israelis.
A second-generation Israeli would be considered 'From Israel by paternal country of origin" in this census, because their paternal country of origin would be Israel.
The bulk of immigration happened in the 1950's-1970's. The number of Israeli's who's ancestors lived in Israel before the establishment of the Israeli state isn't ~~a known or studied figure~~ a definitively answered question, but it's reasonable to assume that it's a minority given the large migrations that happened during and after the Nakba.
The only real information we have regarding the make-up of Palestine before its partitioning are a couple of censuses done during the British occupation, but it was during a period of time when zionist jews were already beginning to migrate. Here's the topline: